Masatoshi Matsumasa and Minoru Murai were able to measure the energetic costs of fiddler crab claws in action by tracking changes in blood glucose (a sugar used to power activity) and blood lactate (a chemical by-product of metabolism and an indication of energy burned) as animals performed various behaviors. By measuring baseline lactate levels in resting animals, and comparing this to the elevated levels they detected during fights, Matsumasa and Murai showed that the energetic cost of waving a claw was substantial.

Emlen, Douglas J, Animal Weapons_ The Evoluti…, loc. 767